Correia-Caeiro C., Kuchenbuch P.H., Oña L.S., Wegdell F., Leroux M., Schuele A., Taglialatela J., Townsend S.W., Surbeck M., Waller B.M., Liebal K. (2025) Adapting the facial action coding system for chimpanzees (Pan troglodytes) to bonobos (Pan paniscus): the ChimpFACS extension for bonobos. PeerJ 13, e19484.
https://doi.org/10.7717/peerj.19484
Jorgewich-Cohen G., Scholte Lubberink N., Wheatley M., Leroux M., Ferrara C.R., Pacciullio Gaspar L., Praschag P., Sánchez-Villagra M.R. (2025) Ontogenetic Changes in Turtle Vocal Behavior. Chelonian Conservation and Biology 24, 46–55.
https://doi.org/10.2744/CCB-1659.1
Leroux M., Lahiff N.J., Zulberti C., Epping A., Uerling C., Taglialatela J.P., Mueller J.L., Watson S.K., Townsend S.W. (2025) Non-adjacent dependency processing (or lack thereof) in bonobos: an artificial grammar experiment. Royal Society Open Science 12, 242173.
https://doi.org/10.1098/rsos.242173
Berthet, M.*, Leroux, M.*, Townsend, S.W., & Watson, S.K. (2025) Methods of examining the processing and production of rule-based sequences in animals. L. Raviv & C. Boeckx (Eds.), The Oxford Handbook of Approaches to Language Evolution. Oxford University Press. *shared first-authorship.
Salis A., Badaire F., Coye C., Leroux M., Lengagne T., Schlenker P., Chemla, E. (2024) Mechanisms of mobbing call recognition: exploring featural decoding in great tits. Animal Behaviour 216, 63-71.
https://doi.org/10.1016/j.anbehav.2024.07.020
Schlenker, P., Salis, A., Badaire, F., Leroux, M., Coye, C & Chemla, E. (2024) Minimal Compositionality versus Bird Implicatures: two theories of ABC-D sequences in Japanese tits. Biological Reviews 99, 1278-1297.
doi.org/10.1111/brv.13068
Leroux, M. (2023) To what extent are call combinations in chimpanzees comparable to syntax in humans? Invited contribution in Revue de Primatologie 14.
doi.org/10.4000/primatologie.16469
Leroux, M., Schel, A.M., Wilke, C., Chandia, B., Zuberbühler, K., Slocombe, K.E., Townsend, S.W. (2023) Call combinations and compositional processing in wild chimpanzees. Nature Communications 14: 2225.
doi.org/10.1038/s41467-023-37816-y
Schlenker, P., Coye, C., Leroux, M., & Chemla, E. (2023) The ABC-D of Animal Linguistics: Are Syntax and Compositionality for Real? Biological Reviews 98, 1142-1159.
doi.org/10.1111/brv.12944
Leroux, M., Chandia, B., Bosshard, A.B., Zuberbühler, K., Townsend, S.W. (2022) Call combinations in chimpanzees: a social tool? Behavioral Ecology 33, 1036-1043.
doi.org/10.1093/beheco/arac074
Bosshard, A.B., Leroux, M., Lester, N.A., Bickel, B., Stoll, S., Townsend, S.W. (2022) From collocations to call-ocations: using linguistic methods to quantify animal call combinations. Behavioral Ecology and Sociobiology 76, 122
doi.org/10.1007/s00265-022-03224-3
Pougnault, L., Levréro, L., Leroux, M., Paulet, J., Bombani, P., Dentressangle, F., Deruti, L., Mulot, B., Lemasson, A. (2022) Social pressure drives “conversational rules” in great apes. Biological Reviews 97, 749-765.
doi.org/10.1111/brv.12821
Leroux, M., Bosshard, A.B., Chandia, B., Manser, A., Zuberbühler, K., Townsend, S.W. (2021) Chimpanzees combine pant-hoots with food-calls into larger structures. Animal Behaviour 179, 41-50.
doi.org/10.1016/j.anbehav.2021.06.026
Leroux, M., Monday, G., Chandia, B., Akankwasa, J.W., Zuberbühler, K., Hobaiter, C., Crockford, C., Townsend, S.W., Asiimwe C., Fedurek, P. (2021) First observation of a chimpanzee with albinism in the wild: Social interactions and subsequent infanticide. American Journal of Primatology e23305.
doi.org/10.1002/ajp.23305
Leroux, M., Gutierrez Al-Khudhairy, O., Perony, N., Townsend, S.W. (2021) Chimpanzee voice prints? Insights from transfer learning experiments from human voices. arXiv 2112.08165.
arXiv:2112.08165v1
Leroux, M., & Townsend, S.W. (2020) Call combinations in great apes and the evolution of syntax. Animal Behaviour and Cognition 7 (2), 131-139.
doi.org/10.26451/abc.07.02.07.2020
Leroux, M., Hetem, R., Hausberger, M., Lemasson, A. (2018) Cheetahs discriminate familiar and unfamiliar human voices. Scientific Reports 8: 15516.
doi.org/10.1038/s41598-018-33971-1
Stomp, M., Leroux, M., Cellier, M., Henry, S., Lemasson, A., Hausberger, M. (2018) An unexpected acoustic indicator of positive emotions in horses. Plos One, 13(7): e0197898.
doi.org/10.1371/journal.pone.0197898
Stomp, M., Leroux, M., Cellier, M., Henry, S., Hausberger, M., Lemasson, A. (2018) Snort acoustic structure codes for positive emotions in horses. The Science of Nature, 105:57.
doi.org/10.1007/s00114-018-1582-9
